The Sun is cutting about 20 production jobs, mainly in print, as it shifts resources to digital.
Nine subeditors on news and a further three on sport have been told they are being made redundant, with a number of other backbench staff across news and features, as well as areas such as design, also being told they are being laid off.
At the same time the title is understood to be looking to hire digital staff, with senior sources saying the layoffs are designed to make __more efficient use of resources.
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The Sun remains the UK’s most popular daily, selling an average of more than 1.7m copies a day in August. However, like most of the rest of the print market it is suffering from a long term shift away from print.
In contrast, the Sun website has been growing rapidly since it dropped its paywall last November. Last month its audience was up 14% to almost 3 million unique browsers a day, almost double what it had a year ago when it was still behind a paywall. It relaunched in June, and has also invested heavily in online betting platform Sun Bets.
However, it still has some way to go to catch up with rivals, with the Independent, Telegraph, Mirror, Guardian and Daily Mail all still significantly ahead online.